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Calendar Name: seminars
Scheduled for: Friday, February 10 2012, 13:30 - 15:00
Event text:    Prof Karin Krupinska
	       
	       Institute of Botany
	       University of Kiel
	       
	       Kiel
	       GERMANY
Details:       " Involvement of the Whirly proteins in communication
	       between the organelles and the nucleus within the plant
	       cell"
	       
	       ABSTRACT
	       Besides a nuclear genome and a mitochondrial genome
	       plant cells have a third genome in their plastids. For
	       coordination of the activities of the three genomes the
	       DNA containing compartments need to exchange
	       information. Whirly proteins form a small family of
	       nucleic acid binding proteins with two to three members
	       exclusively found in higher plants. Investigations on
	       their subcellular localization and their effects on gene
	       expression indicate that the Whirly proteins play a role
	       in transduction of information between organelles and
	       nucleus.
	       After a general introduction on intracellular
	       communication recent results on Whirly1 will be
	       presented. Whirly1 has been identified originally as a
	       nuclear activator of pathogen response genes. Later it
	       was shown that Whirly1 is targeted to plastids and that
	       it has the same molecular weight in plastids and the
	       nucleus, respectively. To investigate whether plastid
	       located Whirly1 might be translocated to the nucleus, we
	       constructed transplastomic tobacco plants synthesizing
	       an HA-tagged version of Whirly1 inside the plastid.
	       Immunological analyses revealed that the tagged protein
	       is also contained in nuclei. Our results give support
	       for a retrograde protein translocation pathway from
	       plastids to the nucleus.

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